Probably. The federal government has the abilility to get access to records that are ordinarily sealed. This is especially true in cases where a security clearance are involved. Since most AF jobs require some level of clearance they will most likely find it even if you lie about it and manage to get in the service. The consequences of the military discharging you for fraudulent enlistment are not good.
I have heard there has been some relaxation of the recuiting standards, but I would be surprised to find you eligible for service in the AF or marines. Your background is always available to scrutiny and as far as I am aware there will not likely be a "sealing" of a DUI charge. Your application requires full disclosure, and as earlier stated, there are serious consequences for fraudulent statements. The services are not granting as many waivers these days. There are other ways of service to your country and I applaude your interest. I sit-down with your local recruiter should inform you more directly.